Effects of Controlling Water to Promote Root and Sparse Sowing to Breed Long-Leaf Age Strong Root Seedlings on Accumulation of Dry Matter and Nitrogen, Phosphorus and Potassium in Flue-Cured Tobacco

doi:10.11924/j.issn.1000-6850.casb2024-0048

Abstract

Abstract:

To explore the effects of cultivating strong root seedlings by controlling water to promote root and sparse sowing to breed long-leaf age seedling on tobacco dry matter and nitrogen, phosphorus and potassium accumulation, two factors of controlling water to promote root and raising long-leaf tobacco seedlings by sparse sowing were designed by using 'Yunyan 87' as material. The dry matter, N, P, K accumulation and distribution, and N, P, K fertilizer utilization efficiency of flue-cured tobacco in the rosette stage, topping stage and round top stage were analyzed, and the η² value was used to evaluate the contribution rate of controlling water to promote root and sparse sowing to breed long-aged strong root seedlings and their interaction. The results showed that increasing the aperture of nursery site, adding microbial agents in the substrate, moisture management adopted by floating seedling at early seedling stage and wet seedling at late seedling stage could improve the dry matter accumulation of tobacco plant, the dry matter distribution of tobacco leaf, the accumulation of N, P and K, the dry matter accumulation efficiency of flue-cured tobacco, the leaf production efficiency, and promote the fertilizer absorption. The contribution rates of controlling water to promote root, sparse sowing to breed long-aged and their interaction on dry matter accumulation of flue-cured tobacco were 30.42%, 51.71% and 17.87%, and those of nitrogen accumulation in flue-cured tobacco were 24.44%, 47.72% and 27.84%, respectively. The contribution rates of phosphorus accumulation in flue-cured tobacco were 25.69%, 45.66% and 28.65%, and those of potassium accumulation in flue-cured tobacco were 30.00%, 44.03% and 25.97%, respectively. Therefore, in the rainy paddy-tobacco growing area of the south, the ways of controlling water to promote root could replace the two-stage seedling cultivation and cultivate strong seedlings with long suitable planting period, which was beneficial to promote early growth and rapid development of flue-cured tobacco and lay the foundation for high quality and high yield.

Key words: control water to promote root, sparse sowing, long-leaf age strong root seedlings, accumulation of tobacco dry matter, accumulation of nitrogen, phosphorus and potassium, flue-cured tobacco production

XIAO Zhipeng, LI Jianhua, WANG Weimin, WEN Weikang, JIANG Zhimin, XU Zhiqiang, MU Tingting, ZENG Huiyu, AN Ran, HUANG Qionghui, DENG Yongsheng, DENG Xiaohua. Effects of Controlling Water to Promote Root and Sparse Sowing to Breed Long-Leaf Age Strong Root Seedlings on Accumulation of Dry Matter and Nitrogen, Phosphorus and Potassium in Flue-Cured Tobacco[J]. Chinese Agricultural Science Bulletin, 2024, 40(27): 27-39.

Figures/Tables 7

References 28

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取样时期	处理	根/（g/株）	茎/（g/株）	叶/（g/株）	干物质积累总量/（g/株）	根占比/%	茎占比/%	叶占比/%
团棵期	A1	0.66±0.17b	2.28±0.61b	7.50±1.65b	10.44±0.27b	6.27±0.92a	21.85±3.01a	71.88±2.69a
	A2	0.73±0.20ab	2.42±0.85a	8.62±1.48a	11.78±0.49a	6.18±0.85a	19.99±3.49a	73.83±3.89a
	A3	0.80±0.29a	2.50±0.81a	8.35±1.04ab	11.65±0.09a	6.68±1.59a	20.93±3.61a	72.39±5.00a
	B1	0.48±0.11b	1.50±0.18b	6.39±0.81b	8.37±0.70b	5.73±1.25b	18.17±3.58	76.09±4.08a
	B2	0.80±0.14a	2.74±0.46a	9.11±0.41a	12.64±0.76a	6.33±1.14ab	21.53±2.77a	72.13±2.85a
	B3	0.91±0.10a	2.96±0.21a	8.98±0.58a	12.85±0.81a	7.06±0.60a	23.06±0.90a	69.88±1.16a
	A1B1	0.47±0.14c	1.70±0.03c	5.40±0.42d	7.57±0.53c	6.16±1.40a	22.51±1.98a	71.33±0.58b
	A1B2	0.70±0.09b	2.37±0.78b	8.78±0.12ab	11.85±0.81b	5.91±1.20b	19.87±5.23a	74.23±4.03ab
	A1B3	0.80±0.01b	2.76±0.27b	8.33±0.20b	11.89±0.49b	6.76±0.16ab	23.17±1.36a	70.07±1.20b
	A2B1	0.51±0.10c	1.34±0.11c	6.73±0.33c	8.57±0.13c	5.91±1.20b	15.62±1.48a	78.47±2.68a
	A2B2	0.76±0.07b	2.90±0.21ab	9.57±0.27a	13.23±0.41a	5.77±0.74b	21.92±0.93a	72.31±0.19b
	A2B3	0.93±0.06ab	3.03±0.07ab	9.57±0.26a	13.53±0.13a	6.85±0.51ab	22.43±0.73a	70.72±1.24b
	A3B1	0.46±0.18c	1.47±0.14c	7.03±0.03c	8.96±0.28c	5.14±1.80b	16.38±1.00a	78.48±2.80a
	A3B2	0.94±0.16ab	2.93±0.13ab	8.98±0.30ab	12.85±0.01b	7.32±1.23ab	22.82±1.03a	69.86±2.26b
	A3B3	0.99±0.10a	3.10±0.12a	9.05±0.19ab	13.15±0.21a	7.57±0.86a	23.59±0.53a	68.84±0.33b
	η²_A	0.355	0.164	0.872	0.816	0.07	0.21	0.21
	η²_B	0.841	0.903	0.979	0.981	0.321	0.658	0.658
	η²_A×B	0.274	0.388	0.653	0.235	0.263	0.594	0.594
打顶期	A1	24.77±1.05b	30.63±1.36b	72.62±1.88b	128.02±3.96b	19.43±1.00a	23.75±1.64a	56.83±1.56a
	A2	25.68±1.16ab	32.81±1.97ab	81.50±1.98a	139.99±3.91a	18.35±0.71a	23.32±2.17a	58.33±2.64a
	A3	26.63±1.40a	33.68±1.05a	78.24±1.92a	138.56±3.61a	19.22±0.39a	24.20±1.99a	56.58±2.22a
	B1	21.77±0.89b	25.01±2.41c	66.96±1.09c	113.73±3.79c	19.20±1.26a	22.03±.002a	58.77±2.98a
	B2	26.83±0.98a	34.61±2.95b	80.10±2.75b	141.54±2.33b	18.97±0.59a	24.42±1.34a	56.61±1.36a
	B3	28.48±1.32a	37.50±1.29a	85.30±1.80a	151.28±2.57a	18.84±0.65a	24.81±0.78a	56.35±1.24a
	A1B1	21.00±0.99c	23.00±0.30c	60.37±1.71d	104.37±3.42d	20.15±1.61a	22.05±1.01a	57.80±2.62a
	A1B2	25.95±0.92b	31.98±1.00b	77.09±1.06b	135.02±1.14c	19.22±0.84a	23.68±0.54a	57.10±0.30a
	A1B3	27.35±0.21ab	36.92±1.39ab	80.41±1.83b	144.68±3.43b	18.91±0.30a	25.52±0.35a	55.58±0.05a
	A2B1	21.80±0.85c	25.70±3.22c	72.07±1.27c	119.57±2.20d	18.24±1.05a	21.52±3.09a	60.23±4.14a
	A2B2	26.70±0.57ab	35.57±3.53ab	82.91±1.40b	145.18±2.69b	18.39±0.05a	24.48±1.98a	57.13±2.02a
	A2B3	28.55±0.35ab	37.15±1.29a	89.52±1.15a	155.22±2.08a	18.42±1.19a	23.95±0.42a	57.63±1.60a
	A3B1	22.50±0.14c	26.34±2.43c	68.43±1.82c	117.27±2.24d	19.20±0.82a	22.51±2.89a	58.29±3.70a
	A3B2	27.85±0.07ab	36.28±2.96ab	80.31±0.81b	144.44±2.22b	19.28±0.25a	25.10±1.66a	55.61±1.41a
	A3B3	29.55±1.91a	38.43±1.46a	85.99±5.41ab	153.96±2.78a	19.19±0.15a	24.97±0.47a	55.84±0.33a
	η²_A	0.609	0.401	0.553	0.717	0.371	0.081	0.188
	η²_B	0.956	0.92	0.846	0.958	0.056	0.508	0.313
	η²_A×B	0.045	0.135	0.092	0.077	0.166	0.109	0.076
圆顶期	A1	63.13±2.22b	74.84±2.13b	114.95±2.52b	252.92±3.78b	24.99±0.95a	29.64±0.98a	45.37±1.74a
	A2	67.13±2.44a	77.95±2.75a	125.63±2.78a	270.70±2.44a	24.85±0.68a	28.82±0.94a	46.32±1.22a
	A3	68.55±2.34a	78.82±2.33a	121.44±2.81a	268.80±2.77a	25.58±1.11a	29.37±0.74a	45.06±1.68a
	B1	59.59±2.05b	68.91±2.45c	103.22±3.56c	231.72±2.10c	25.70±0.96a	29.75±0.38a	44.56±0.83a
	B2	67.04±3.24a	76.94±2.86b	121.04±2.73b	265.02±3.03b	25.31±0.98a	29.08±1.47a	45.61±2.26a
	B3	72.18±2.47a	85.74±2.44a	137.76±2.50a	295.68±2.19a	24.41±0.19a	29.00±0.32a	46.58±0.36a
	A1B1	55.13±2.77c	66.00±0.87c	99.75±1.80d	220.87±0.10c	24.96±1.24a	29.88±0.41a	45.16±0.84a
	A1B2	63.58±2.25b	74.68±3.22b	110.80±2.87c	249.06±2.40b	25.54±1.35a	30.00±1.82a	44.46±3.18a
	A1B3	70.69±2.26ab	83.83±0.96ab	134.30±4.29ab	288.82±2.50ab	24.47±0.15a	29.03±0.42a	46.50±0.28a
	A2B1	60.96±1.39bc	70.15±1.88b	106.83±2.64cd	237.94±2.90b	25.62±0.05a	29.48±0.06a	44.90±0.01a
	A2B2	67.97±2.22ab	77.07±3.05ab	129.65±4.93ab	274.69±4.09ab	24.74±0.44a	28.07±1.53a	47.19±1.09a
	A2B3	72.45±2.67a	86.62±3.69a	140.41±2.14a	299.48±2.51a	24.19±0.12a	28.92±0.02a	46.88±0.09a
	A3B1	62.68±1.12b	70.60±0.09b	103.08±1.68cd	236.35±4.89b	26.51±0.77a	29.87±0.58a	43.62±0.19a
	A3B2	69.56±2.05ab	79.08±1.34ab	122.67±2.54b	271.31±2.14ab	25.65±1.34a	29.16±1.16a	45.18±2.49a
	A3B3	73.40±1.27a	86.78±2.08a	138.56±2.48a	298.74±2.83a	24.57±0.04a	29.06±0.55a	46.37±0.51a
	η²_A	0.626	0.541	0.507	0.692	0.229	0.204	0.219
	η²_B	0.895	0.95	0.914	0.96	0.465	0.2	0.399
	η²_A×B	0.199	0.073	0.238	0.201	0.229	0.195	0.238

取样时期	处理	根/（g/株）	茎/（g/株）	叶/（g/株）	干物质积累总量/（g/株）	根占比/%	茎占比/%	叶占比/%
团棵期	A1	0.66±0.17b	2.28±0.61b	7.50±1.65b	10.44±0.27b	6.27±0.92a	21.85±3.01a	71.88±2.69a
	A2	0.73±0.20ab	2.42±0.85a	8.62±1.48a	11.78±0.49a	6.18±0.85a	19.99±3.49a	73.83±3.89a
	A3	0.80±0.29a	2.50±0.81a	8.35±1.04ab	11.65±0.09a	6.68±1.59a	20.93±3.61a	72.39±5.00a
	B1	0.48±0.11b	1.50±0.18b	6.39±0.81b	8.37±0.70b	5.73±1.25b	18.17±3.58	76.09±4.08a
	B2	0.80±0.14a	2.74±0.46a	9.11±0.41a	12.64±0.76a	6.33±1.14ab	21.53±2.77a	72.13±2.85a
	B3	0.91±0.10a	2.96±0.21a	8.98±0.58a	12.85±0.81a	7.06±0.60a	23.06±0.90a	69.88±1.16a
	A1B1	0.47±0.14c	1.70±0.03c	5.40±0.42d	7.57±0.53c	6.16±1.40a	22.51±1.98a	71.33±0.58b
	A1B2	0.70±0.09b	2.37±0.78b	8.78±0.12ab	11.85±0.81b	5.91±1.20b	19.87±5.23a	74.23±4.03ab
	A1B3	0.80±0.01b	2.76±0.27b	8.33±0.20b	11.89±0.49b	6.76±0.16ab	23.17±1.36a	70.07±1.20b
	A2B1	0.51±0.10c	1.34±0.11c	6.73±0.33c	8.57±0.13c	5.91±1.20b	15.62±1.48a	78.47±2.68a
	A2B2	0.76±0.07b	2.90±0.21ab	9.57±0.27a	13.23±0.41a	5.77±0.74b	21.92±0.93a	72.31±0.19b
	A2B3	0.93±0.06ab	3.03±0.07ab	9.57±0.26a	13.53±0.13a	6.85±0.51ab	22.43±0.73a	70.72±1.24b
	A3B1	0.46±0.18c	1.47±0.14c	7.03±0.03c	8.96±0.28c	5.14±1.80b	16.38±1.00a	78.48±2.80a
	A3B2	0.94±0.16ab	2.93±0.13ab	8.98±0.30ab	12.85±0.01b	7.32±1.23ab	22.82±1.03a	69.86±2.26b
	A3B3	0.99±0.10a	3.10±0.12a	9.05±0.19ab	13.15±0.21a	7.57±0.86a	23.59±0.53a	68.84±0.33b
	η²_A	0.355	0.164	0.872	0.816	0.07	0.21	0.21
	η²_B	0.841	0.903	0.979	0.981	0.321	0.658	0.658
	η²_A×B	0.274	0.388	0.653	0.235	0.263	0.594	0.594
打顶期	A1	24.77±1.05b	30.63±1.36b	72.62±1.88b	128.02±3.96b	19.43±1.00a	23.75±1.64a	56.83±1.56a
	A2	25.68±1.16ab	32.81±1.97ab	81.50±1.98a	139.99±3.91a	18.35±0.71a	23.32±2.17a	58.33±2.64a
	A3	26.63±1.40a	33.68±1.05a	78.24±1.92a	138.56±3.61a	19.22±0.39a	24.20±1.99a	56.58±2.22a
	B1	21.77±0.89b	25.01±2.41c	66.96±1.09c	113.73±3.79c	19.20±1.26a	22.03±.002a	58.77±2.98a
	B2	26.83±0.98a	34.61±2.95b	80.10±2.75b	141.54±2.33b	18.97±0.59a	24.42±1.34a	56.61±1.36a
	B3	28.48±1.32a	37.50±1.29a	85.30±1.80a	151.28±2.57a	18.84±0.65a	24.81±0.78a	56.35±1.24a
	A1B1	21.00±0.99c	23.00±0.30c	60.37±1.71d	104.37±3.42d	20.15±1.61a	22.05±1.01a	57.80±2.62a
	A1B2	25.95±0.92b	31.98±1.00b	77.09±1.06b	135.02±1.14c	19.22±0.84a	23.68±0.54a	57.10±0.30a
	A1B3	27.35±0.21ab	36.92±1.39ab	80.41±1.83b	144.68±3.43b	18.91±0.30a	25.52±0.35a	55.58±0.05a
	A2B1	21.80±0.85c	25.70±3.22c	72.07±1.27c	119.57±2.20d	18.24±1.05a	21.52±3.09a	60.23±4.14a
	A2B2	26.70±0.57ab	35.57±3.53ab	82.91±1.40b	145.18±2.69b	18.39±0.05a	24.48±1.98a	57.13±2.02a
	A2B3	28.55±0.35ab	37.15±1.29a	89.52±1.15a	155.22±2.08a	18.42±1.19a	23.95±0.42a	57.63±1.60a
	A3B1	22.50±0.14c	26.34±2.43c	68.43±1.82c	117.27±2.24d	19.20±0.82a	22.51±2.89a	58.29±3.70a
	A3B2	27.85±0.07ab	36.28±2.96ab	80.31±0.81b	144.44±2.22b	19.28±0.25a	25.10±1.66a	55.61±1.41a
	A3B3	29.55±1.91a	38.43±1.46a	85.99±5.41ab	153.96±2.78a	19.19±0.15a	24.97±0.47a	55.84±0.33a
	η²_A	0.609	0.401	0.553	0.717	0.371	0.081	0.188
	η²_B	0.956	0.92	0.846	0.958	0.056	0.508	0.313
	η²_A×B	0.045	0.135	0.092	0.077	0.166	0.109	0.076
圆顶期	A1	63.13±2.22b	74.84±2.13b	114.95±2.52b	252.92±3.78b	24.99±0.95a	29.64±0.98a	45.37±1.74a
	A2	67.13±2.44a	77.95±2.75a	125.63±2.78a	270.70±2.44a	24.85±0.68a	28.82±0.94a	46.32±1.22a
	A3	68.55±2.34a	78.82±2.33a	121.44±2.81a	268.80±2.77a	25.58±1.11a	29.37±0.74a	45.06±1.68a
	B1	59.59±2.05b	68.91±2.45c	103.22±3.56c	231.72±2.10c	25.70±0.96a	29.75±0.38a	44.56±0.83a
	B2	67.04±3.24a	76.94±2.86b	121.04±2.73b	265.02±3.03b	25.31±0.98a	29.08±1.47a	45.61±2.26a
	B3	72.18±2.47a	85.74±2.44a	137.76±2.50a	295.68±2.19a	24.41±0.19a	29.00±0.32a	46.58±0.36a
	A1B1	55.13±2.77c	66.00±0.87c	99.75±1.80d	220.87±0.10c	24.96±1.24a	29.88±0.41a	45.16±0.84a
	A1B2	63.58±2.25b	74.68±3.22b	110.80±2.87c	249.06±2.40b	25.54±1.35a	30.00±1.82a	44.46±3.18a
	A1B3	70.69±2.26ab	83.83±0.96ab	134.30±4.29ab	288.82±2.50ab	24.47±0.15a	29.03±0.42a	46.50±0.28a
	A2B1	60.96±1.39bc	70.15±1.88b	106.83±2.64cd	237.94±2.90b	25.62±0.05a	29.48±0.06a	44.90±0.01a
	A2B2	67.97±2.22ab	77.07±3.05ab	129.65±4.93ab	274.69±4.09ab	24.74±0.44a	28.07±1.53a	47.19±1.09a
	A2B3	72.45±2.67a	86.62±3.69a	140.41±2.14a	299.48±2.51a	24.19±0.12a	28.92±0.02a	46.88±0.09a
	A3B1	62.68±1.12b	70.60±0.09b	103.08±1.68cd	236.35±4.89b	26.51±0.77a	29.87±0.58a	43.62±0.19a
	A3B2	69.56±2.05ab	79.08±1.34ab	122.67±2.54b	271.31±2.14ab	25.65±1.34a	29.16±1.16a	45.18±2.49a
	A3B3	73.40±1.27a	86.78±2.08a	138.56±2.48a	298.74±2.83a	24.57±0.04a	29.06±0.55a	46.37±0.51a
	η²_A	0.626	0.541	0.507	0.692	0.229	0.204	0.219
	η²_B	0.895	0.95	0.914	0.96	0.465	0.2	0.399
	η²_A×B	0.199	0.073	0.238	0.201	0.229	0.195	0.238

取样时期	处理	根/（mg/株）	茎/（mg/株）	叶/（mg/株）	氮积累总量（/ mg/株）	根占比/%	茎占比/%	叶占比/%
团棵期	A1	16.80±4.57c	103.30±4.82c	211.01±6.23c	331.12±11.46b	5.17±1.34a	30.76±8.12a	64.07±8.70a
	A2	19.54±2.67b	110.96±4.18b	323.75±13.33b	454.25±18.65a	4.38±1.05a	25.85±2.75a	69.76±2.68a
	A3	22.32±2.61a	134.21±6.11a	296.04±9.44a	452.57±28.07a	4.82±1.60a	29.62±7.95a	65.56±9.40a
	B1	11.18±1.94c	63.32±1.27c	231.65±3.07c	306.16±13.46c	3.69±0.81a	20.80±3.53b	75.50±3.75a
	B2	21.32±4.69b	138.30±3.57b	290.03±3.80b	449.64±16.57b	6.14±1.00a	39.37±4.80a	54.48±4.41b
	B3	26.17±4.93a	146.85±3.17a	409.13±11.82a	582.14±11.99a	4.54±0.69a	26.05±5.24b	69.4±5.63a
	A1B1	11.62±3.40c	59.98±0.64c	192.23±5.85c	263.82±9.89f	4.38±1.13a	22.75±0.61b	72.87±0.51ab
	A1B2	18.34±1.40b	109.35±3.97b	155.18±3.91c	282.87±4.65e	6.56±1.30a	38.08±9.46a	55.36±8.16c
	A1B3	20.45±2.40b	140.58±11.23ab	285.64±3.35b	446.67±1.28c	4.57±0.43a	31.45±1.79ab	63.97±2.22b
	A2B1	11.49±0.25c	54.32±8.08c	258.74±4.43b	324.56±6.59d	3.54±0.01a	16.77±2.83b	79.69±2.83a
	A2B2	19.02±3.12b	138.97±5.10ab	182.70±0.43c	340.69±1.55d	5.58±0.94a	40.79±1.31a	53.63±0.37c
	A2B3	28.12±2.71a	139.58±6.76ab	529.81±1.31a	697.50±1.78a	4.03±0.33a	20.01±0.66b	75.97±0.99ab
	A3B1	10.44±2.35c	75.67±8.93c	243.98±4.34b	330.09±6.94d	3.15±0.64a	22.90±2.22b	73.94±2.87ab
	A3B2	26.59±3.82ab	166.57±9.25a	232.21±3.33b	425.36±2.26c	6.28±1.20a	39.26±4.04a	54.47±5.24c
	A3B3	29.93±2.68a	160.40±2.02a	411.93±3.14ab	602.26±72.48b	5.03±1.05a	26.71±1.22ab	68.26±2.27ab
	η²_A	0.589	0.589	0.885	0.892	0.209	0.385	0.462
	η²_B	0.917	0.921	0.968	0.973	0.724	0.897	0.92
	η²_A×B	0.6	0.357	0.844	0.806	0.196	0.465	0.451
打顶期	A1	771.49±19.51a	656.91±36.86b	1499.94±39.57b	2928.34±83.69b	26.84±3.76a	21.13±6.44a	52.03±5.36a
	A2	791.91±28.19a	893.24±42.63a	1869.50±49.79a	3554.65±39.67a	22.31±2.12a	24.05±5.09a	53.65±4.79a
	A3	781.22±24.57a	869.76±40.86a	1770.35±32.72a	3421.33±79.57a	22.73±3.99a	24.38±7.00a	52.89±7.83a
	B1	521.18±52.25c	386.03±13.93c	1332.05±25.81c	2239.27±29.39c	23.61±3.80a	17.09±3.24b	59.30±2.04a
	B2	852.74±12.76b	1195.58±26.72a	2008.69±39.53a	4057.01±63.92a	21.36±3.73a	29.26±4.12a	49.37±4.80b
	B3	970.69±17.15a	838.30±18.21b	1799.04±15.71b	3608.04±19.82b	26.91±1.60a	23.21±2.74ab	49.89±3.52b
	A1B1	553.34±19.47c	259.22±1.45c	1134.67±28.37c	1947.23±36.39d	28.45±0.39a	13.57±2.70b	57.97±3.09b
	A1B2	797.37±8.23b	935.41±23.54b	1678.24±42.11b	3411.03±66.41b	23.97±6.70a	27.23±2.12ab	48.79±4.58b
	A1B3	963.75±8.81ab	776.09±15.4b	1686.91±25.64b	3426.76±29.85b	28.09±1.00a	22.60±1.56ab	49.31±2.56b
	A2B1	501.10±2.43c	471.02±6.80c	1390.48±6.45c	2362.60±24.81c	21.24±1.23a	19.90±1.56b	58.86±0.34ab
	A2B2	969.77±4.19ab	1395.99±20.42a	2305.75±14.85a	4671.51±18.75a	20.76±0.17a	29.81±3.86a	49.43±4.03b
	A2B3	904.87±3.6ab	812.72±6.37b	1912.27±10.35ab	3629.86±11.58b	24.93±0.12a	22.43±2.41ab	52.64±2.52ab
	A3B1	509.12±4.23c	427.84±8.93c	1471.02±8.57c	2407.98±9.87c	21.14±0.16a	17.79±1.09b	61.07±0.93a
	A3B2	791.10±3.68b	1255.33±20.52a	2042.08±8.32ab	4088.50±8.12ab	19.36±1.54a	30.75±7.22a	49.90±8.76b
	A3B3	1043.45±9.58a	926.10±17.36b	1797.96±8.79b	3767.51±4.02b	27.70±0.21a	24.60±4.85ab	47.70±4.85b
	η²_A	0.037	0.461	0.461	0.674	0.6	0.251	0.047
	η²_B	0.952	0.893	0.737	0.945	0.651	0.796	0.7
	η²_A×B	0.657	0.284	0.205	0.511	0.371	0.163	0.152
圆顶期	A1	868.41±81.60c	1110.05±0.64b	2950.42±21.92a	4928.87±30.96c	17.65±2.12a	22.53±0.59b	59.82±2.71a
	A2	1098.71±72.54b	1127.14±32.85b	3056.28±37.58a	5282.12±72.20b	20.86±2.45a	21.38±1.72b	57.75±4.17b
	A3	1258.63±85.1a	1786.49±37.97a	3022.56±56.56a	6067.68±79.63a	20.81±1.10a	29.52±1.28a	49.67±2.38b
	B1	1075.25±86.06b	1341.23±50.77b	3009.75±31.92c	5426.23±69.73c	19.77±2.25a	24.48±4.06ab	55.75±5.39b
	B2	851.68±29.96b	1695.28±38.02a	3254.24±91.69b	5801.20±69.67b	14.57±2.01a	29.37±2.58a	56.06±0.57b
	B3	1393.38±67.12a	1331.91±93.61b	4980.52±84.62a	7705.82±58.13a	18.09±0.20a	17.35±2.25b	64.56±2.45a
	A1B1	1158.56±19.72b	1756.75±20.73a	2871.16±73.61c	5786.47±24.06ab	20.03±0.4a	30.34±0.96a	49.63±0.56c
	A1B2	1134.54±62.36b	1594.65±27.10b	3701.98±43.50ab	6431.16±52.55a	17.56±2.64a	25.69±6.66b	56.75±6.80b
	A1B3	864.36±45.41c	1560.66±88.10b	3442.40±68.91ab	5867.41±92.41ab	14.58±1.82a	26.83±2.85b	58.59±1.04b
	A2B1	1110.63±65.91b	1174.05±24.72c	3861.14±83.57a	6145.83±7.06ab	18.07±1.05a	19.10±2.01c	62.83±3.06a
	A2B2	1313.78±97.00a	1730.13±73.64a	2731.68±84.16c	5775.59±60.8ab	22.78±2.31a	29.95±0.44a	47.27±1.87c
	A2B3	1096.25±35.19	1488.28±65.96b	3345.07±52.41ab	5929.61±45.12ab	18.48±3.93a	25.29±5.24b	56.23±7.38b
	A3B1	861.48±49.18c	1455.33±77.48b	3215.68±46.91b	5532.49±67.26b	15.60±2.21a	26.24±3.55b	58.16±2.16b
	A3B2	1200.91±58.36a	1211.03±19.53c	3965.98±90.69a	6377.92±24.27a	19.01±1.87a	19.28±2.38c	61.71±4.07a
	A3B3	1243.66±91.44a	1757.79±91.89a	2875.13±75.89c	5876.58±77.81ab	21.21±1.72a	29.94±0.83a	48.86±1.84c
	η²_A	0.071	0.725	0.527	0.577	0.365	0.132	0.056
	η²_B	0.787	0.924	0.744	0.494	0.788	0.921	0.912
	η²_A×B	0.469	0.758	0.671	0.655	0.385	0.668	0.576

取样时期	处理	根/（mg/株）	茎/（mg/株）	叶/（mg/株）	氮积累总量（/ mg/株）	根占比/%	茎占比/%	叶占比/%
团棵期	A1	16.80±4.57c	103.30±4.82c	211.01±6.23c	331.12±11.46b	5.17±1.34a	30.76±8.12a	64.07±8.70a
	A2	19.54±2.67b	110.96±4.18b	323.75±13.33b	454.25±18.65a	4.38±1.05a	25.85±2.75a	69.76±2.68a
	A3	22.32±2.61a	134.21±6.11a	296.04±9.44a	452.57±28.07a	4.82±1.60a	29.62±7.95a	65.56±9.40a
	B1	11.18±1.94c	63.32±1.27c	231.65±3.07c	306.16±13.46c	3.69±0.81a	20.80±3.53b	75.50±3.75a
	B2	21.32±4.69b	138.30±3.57b	290.03±3.80b	449.64±16.57b	6.14±1.00a	39.37±4.80a	54.48±4.41b
	B3	26.17±4.93a	146.85±3.17a	409.13±11.82a	582.14±11.99a	4.54±0.69a	26.05±5.24b	69.4±5.63a
	A1B1	11.62±3.40c	59.98±0.64c	192.23±5.85c	263.82±9.89f	4.38±1.13a	22.75±0.61b	72.87±0.51ab
	A1B2	18.34±1.40b	109.35±3.97b	155.18±3.91c	282.87±4.65e	6.56±1.30a	38.08±9.46a	55.36±8.16c
	A1B3	20.45±2.40b	140.58±11.23ab	285.64±3.35b	446.67±1.28c	4.57±0.43a	31.45±1.79ab	63.97±2.22b
	A2B1	11.49±0.25c	54.32±8.08c	258.74±4.43b	324.56±6.59d	3.54±0.01a	16.77±2.83b	79.69±2.83a
	A2B2	19.02±3.12b	138.97±5.10ab	182.70±0.43c	340.69±1.55d	5.58±0.94a	40.79±1.31a	53.63±0.37c
	A2B3	28.12±2.71a	139.58±6.76ab	529.81±1.31a	697.50±1.78a	4.03±0.33a	20.01±0.66b	75.97±0.99ab
	A3B1	10.44±2.35c	75.67±8.93c	243.98±4.34b	330.09±6.94d	3.15±0.64a	22.90±2.22b	73.94±2.87ab
	A3B2	26.59±3.82ab	166.57±9.25a	232.21±3.33b	425.36±2.26c	6.28±1.20a	39.26±4.04a	54.47±5.24c
	A3B3	29.93±2.68a	160.40±2.02a	411.93±3.14ab	602.26±72.48b	5.03±1.05a	26.71±1.22ab	68.26±2.27ab
	η²_A	0.589	0.589	0.885	0.892	0.209	0.385	0.462
	η²_B	0.917	0.921	0.968	0.973	0.724	0.897	0.92
	η²_A×B	0.6	0.357	0.844	0.806	0.196	0.465	0.451
打顶期	A1	771.49±19.51a	656.91±36.86b	1499.94±39.57b	2928.34±83.69b	26.84±3.76a	21.13±6.44a	52.03±5.36a
	A2	791.91±28.19a	893.24±42.63a	1869.50±49.79a	3554.65±39.67a	22.31±2.12a	24.05±5.09a	53.65±4.79a
	A3	781.22±24.57a	869.76±40.86a	1770.35±32.72a	3421.33±79.57a	22.73±3.99a	24.38±7.00a	52.89±7.83a
	B1	521.18±52.25c	386.03±13.93c	1332.05±25.81c	2239.27±29.39c	23.61±3.80a	17.09±3.24b	59.30±2.04a
	B2	852.74±12.76b	1195.58±26.72a	2008.69±39.53a	4057.01±63.92a	21.36±3.73a	29.26±4.12a	49.37±4.80b
	B3	970.69±17.15a	838.30±18.21b	1799.04±15.71b	3608.04±19.82b	26.91±1.60a	23.21±2.74ab	49.89±3.52b
	A1B1	553.34±19.47c	259.22±1.45c	1134.67±28.37c	1947.23±36.39d	28.45±0.39a	13.57±2.70b	57.97±3.09b
	A1B2	797.37±8.23b	935.41±23.54b	1678.24±42.11b	3411.03±66.41b	23.97±6.70a	27.23±2.12ab	48.79±4.58b
	A1B3	963.75±8.81ab	776.09±15.4b	1686.91±25.64b	3426.76±29.85b	28.09±1.00a	22.60±1.56ab	49.31±2.56b
	A2B1	501.10±2.43c	471.02±6.80c	1390.48±6.45c	2362.60±24.81c	21.24±1.23a	19.90±1.56b	58.86±0.34ab
	A2B2	969.77±4.19ab	1395.99±20.42a	2305.75±14.85a	4671.51±18.75a	20.76±0.17a	29.81±3.86a	49.43±4.03b
	A2B3	904.87±3.6ab	812.72±6.37b	1912.27±10.35ab	3629.86±11.58b	24.93±0.12a	22.43±2.41ab	52.64±2.52ab
	A3B1	509.12±4.23c	427.84±8.93c	1471.02±8.57c	2407.98±9.87c	21.14±0.16a	17.79±1.09b	61.07±0.93a
	A3B2	791.10±3.68b	1255.33±20.52a	2042.08±8.32ab	4088.50±8.12ab	19.36±1.54a	30.75±7.22a	49.90±8.76b
	A3B3	1043.45±9.58a	926.10±17.36b	1797.96±8.79b	3767.51±4.02b	27.70±0.21a	24.60±4.85ab	47.70±4.85b
	η²_A	0.037	0.461	0.461	0.674	0.6	0.251	0.047
	η²_B	0.952	0.893	0.737	0.945	0.651	0.796	0.7
	η²_A×B	0.657	0.284	0.205	0.511	0.371	0.163	0.152
圆顶期	A1	868.41±81.60c	1110.05±0.64b	2950.42±21.92a	4928.87±30.96c	17.65±2.12a	22.53±0.59b	59.82±2.71a
	A2	1098.71±72.54b	1127.14±32.85b	3056.28±37.58a	5282.12±72.20b	20.86±2.45a	21.38±1.72b	57.75±4.17b
	A3	1258.63±85.1a	1786.49±37.97a	3022.56±56.56a	6067.68±79.63a	20.81±1.10a	29.52±1.28a	49.67±2.38b
	B1	1075.25±86.06b	1341.23±50.77b	3009.75±31.92c	5426.23±69.73c	19.77±2.25a	24.48±4.06ab	55.75±5.39b
	B2	851.68±29.96b	1695.28±38.02a	3254.24±91.69b	5801.20±69.67b	14.57±2.01a	29.37±2.58a	56.06±0.57b
	B3	1393.38±67.12a	1331.91±93.61b	4980.52±84.62a	7705.82±58.13a	18.09±0.20a	17.35±2.25b	64.56±2.45a
	A1B1	1158.56±19.72b	1756.75±20.73a	2871.16±73.61c	5786.47±24.06ab	20.03±0.4a	30.34±0.96a	49.63±0.56c
	A1B2	1134.54±62.36b	1594.65±27.10b	3701.98±43.50ab	6431.16±52.55a	17.56±2.64a	25.69±6.66b	56.75±6.80b
	A1B3	864.36±45.41c	1560.66±88.10b	3442.40±68.91ab	5867.41±92.41ab	14.58±1.82a	26.83±2.85b	58.59±1.04b
	A2B1	1110.63±65.91b	1174.05±24.72c	3861.14±83.57a	6145.83±7.06ab	18.07±1.05a	19.10±2.01c	62.83±3.06a
	A2B2	1313.78±97.00a	1730.13±73.64a	2731.68±84.16c	5775.59±60.8ab	22.78±2.31a	29.95±0.44a	47.27±1.87c
	A2B3	1096.25±35.19	1488.28±65.96b	3345.07±52.41ab	5929.61±45.12ab	18.48±3.93a	25.29±5.24b	56.23±7.38b
	A3B1	861.48±49.18c	1455.33±77.48b	3215.68±46.91b	5532.49±67.26b	15.60±2.21a	26.24±3.55b	58.16±2.16b
	A3B2	1200.91±58.36a	1211.03±19.53c	3965.98±90.69a	6377.92±24.27a	19.01±1.87a	19.28±2.38c	61.71±4.07a
	A3B3	1243.66±91.44a	1757.79±91.89a	2875.13±75.89c	5876.58±77.81ab	21.21±1.72a	29.94±0.83a	48.86±1.84c
	η²_A	0.071	0.725	0.527	0.577	0.365	0.132	0.056
	η²_B	0.787	0.924	0.744	0.494	0.788	0.921	0.912
	η²_A×B	0.469	0.758	0.671	0.655	0.385	0.668	0.576

取样时期	处理	根/（mg/株）	茎/（mg/株）	叶/（mg/株）	磷积累总量/（mg/株）	根占比/%	茎占比/%	叶占比/%
团棵期	A1	1.92±0.71b	7.67±3.00a	30.45±1.61a	40.04±3.52a	4.78±0.73a	19.48±4.55a	75.74±4.11a
	A2	2.20±0.95a	8.16±3.78a	37.83±1.53a	48.19±6.15a	4.48±0.75a	16.08±3.21a	79.44±3.46a
	A3	2.30±1.11a	7.44±2.78a	33.98±2.49a	43.72±5.96a	5.02±1.19a	17.19±2.45a	77.79±2.87a
	B1	1.06±0.27b	4.10±0.84b	19.39±3.07b	25.55±2.54b	4.29±0.99a	17.05±4.81a	78.66±4.93a
	B2	2.42±0.41a	9.12±2.08a	40.04±2.65a	51.57±3.81a	4.72±0.91a	17.55±3.33a	77.73±3.27a
	B3	2.95±0.45a	10.06±1.32a	42.82±2.97a	55.83±5.17a	5.27±0.56a	18.15±3.05a	76.57±2.79a
	A1B1	2.07±0.32bc	4.85±0.92c	16.81±1.53c	23.72±0.93b	4.67±1.21a	21.44±4.91a	73.88±3.70a
	A1B2	2.24±0.39b	7.33±2.85b	37.88±0.07ab	47.45±2.39ab	4.75±1.06a	15.32±5.24b	79.94±4.18a
	A1B3	2.46±0. 10ab	10.83±0.22a	36.65±0.76ab	49.94±1.08ab	4.93±0.10a	21.68±0.03a	73.39±5.07a
	A2B1	2.16±0.15b	3.33±0.31d	23.13±0.72b	28.62±0.26b	4.19±0.59a	12.08±1.24c	83.73±1.83a
	A2B2	2.21±0.31b	10.16±0.86ab	43.20±0.44ab	55.57±0.99ab	3.97±0.63a	18.28±1.22a	77.75±0.60a
	A2B3	3.24±0.26a	10.98±0.18a	47.16±0.14a	61.38±0.23a	5.27±0.40a	17.90±0.36a	76.83±0.05a
	A3B1	1.95±0.45c	4.12±0.53c	18.25±0.64bc	24.31±1.62b	4.01±1.63a	17.63±1.05a	78.36±2.68a
	A3B2	2.81±0.36ab	9.85±1.75ab	39.02±1.89ab	51.68±0.23ab	5.44±0.68a	19.06±3.31a	75.51±3.99a
	A3B3	3.15±0.48a	8.36±0.07ab	44.66±1.47ab	56.17±1.06ab	5.62±0.95a	14.88±0.16b	79.50±1.12a
	η²_A	0.316	0.109	0.943	0.941	0.105	0.351	0.41
	η²_B	0.919	0.903	0.995	0.996	0.278	0.052	0.181
	η²_A×B	0.431	0.589	0.767	0.671	0.225	0.632	0.656
打顶期	A1	58.41±15.74b	48.34±5.26b	230.49±7.17b	337.24±15.45b	17.72±2.52a	14.47±1.78a	67.81±2.89a
	A2	73.91±8.70a	69.05±6.84a	263.57±8.98a	406.54±16.47a	17.76±3.01a	17.38±3.26a	64.86±2.52a
	A3	68.56±4.98a	66.30±9.24a	276.57±6.47a	411.44±9.19a	16.33±3.72a	16.00±2.77a	67.67±5.75a
	B1	40.66±2.96c	41.59±9.96b	183.33±6.09c	265.57±6.39b	16.01±3.82a	15.96±3.48a	68.03±6.00a
	B2	70.13±7.29b	75.83±5.48a	285.81±6.12b	431.77±4.75a	16.25±1.19a	17.44±2.53a	66.31±2.08a
	B3	90.09±7.27a	66.28±10.09a	301.49±8.78a	457.86±7.36a	19.54±2.34a	14.45±1.65a	66.00±3.35a
	A1B1	39.69±5.71c	30.36±0.07d	132.58±5.65c	202.63±10.00c	19.68±3.79a	15.00±0.71a	65.32±4.50a
	A1B2	62.96±6.98b	59.25±3.33b	270.26±2.03ab	392.47±4.32b	16.05±1.95a	15.08±3.23a	68.87±1.28a
	A1B3	72.57±3.71ab	55.41±1.17b	288.63±2.74ab	416.61±3.29ab	17.42±0.08a	13.33±1.03a	69.25±1.11a
	A2B1	42.54±0.26c	51.71±5.10b	177.36±2.22bc	271.60±8.38c	15.69±0.97a	19.15±3.18a	65.16±4.14a
	A2B2	73.56±7.31ab	86.34±8.57a	293.46±1.58ab	453.35±0.33ab	16.22±1.60a	19.05±1.90a	64.73±0.30a
	A2B3	105.63±9.24a	69.12±10.95ab	319.91±5.87a	494.65±4.32a	21.35±1.68a	13.96±2.09a	64.69±3.77a
	A3B1	39.73±0.67c	42.70±3.23c	240.06±8.85b	322.49±6.28c	12.65±2.79a	13.74±4.25a	73.61±7.04a
	A3B2	73.89±3.02ab	81.91±10.17a	293.71±3.81ab	449.51±7.00ab	16.47±0.68a	18.19±0.77a	65.34±0.08a
	A3B3	92.07±6.94a	74.30±2.57ab	295.94±3.13ab	462.31±6.38ab	19.86±2.96a	16.07±0.01a	64.07±2.95a
	η²_A	0.584	0.745	0.45	0.709	0.161	0.344	0.228
	η²_B	0.933	0.878	0.857	0.939	0.53	0.354	0.112
	η²_A×B	0.498	0.243	0.447	0.352	0.581	0.397	0.498
圆顶期	A1	118.88±11.01c	125.79±11.05c	316.23±12.34b	560.90±13.38b	21.20±0.51b	22.41±1.44b	56.39±0.93a
	A2	158.31±11.63b	223.51±8.05b	501.57±67.13a	883.39±86.82a	17.94±0.45b	25.38±1.58b	56.68±2.03a
	A3	211.79±20.17a	258.58±5.81a	323.22±42.3b	793.60±68.28a	26.68±0.25a	32.67±2.08a	40.65±1.83b
	B1	163.00±42.99a	202.63±6.91b	380.34±10.44b	745.96±56.96b	21.94±3.96a	26.82±4.91a	51.24±8.30a
	B2	167.02±15.51a	202.74±7.40b	379.23±15.16b	749.00±23.27b	19.03±1.54a	28.23±1.94a	52.74±0.40a
	B3	163.91±18.25a	268.76±7.10a	531.63±75.81a	964.31±11.16a	16.99±0.11a	27.99±2.20a	55.02±2.09a
	A1B1	231.84±8.14a	242.25±4.08ab	369.17±78.95b	843.25±29.73b	27.53±1.94a	28.83±5.89a	43.64±7.82c
	A1B2	177.59±4.15ab	237.92±3.24ab	426.68±9.24ab	842.19±19.86b	21.18±5.12ab	28.35±2.97ab	50.47±6.49b
	A1B3	127.43±15.25b	164.94±11.75b	414.5±28.87ab	706.87±25.37c	18.08±2.81b	23.32±0.83b	58.60±1.98a
	A2B1	190.13±22.21ab	234.98±23.83ab	559.80±7.05a	984.91±5.43a	19.30±2.15b	23.87±2.55b	56.84±0.40ab
	A2B2	247.64±32.9a	299.93±31.81a	344.66±9.74b	892.24±9.83b	27.72±0.65a	34.02±7.29a	38.26±6.65
	A2B3	188.40±7.04ab	233.29±3.17ab	439.65±18.1ab	861.34±34.47b	21.70±4.96ab	27.07±6.41ab	51.23±10.55ab
	A3B1	127.78±12.67b	164.49±35.32b	369.99±46.87b	662.26±8.41c	19.44±2.04b	24.65±3.02b	55.91±2.83ab
	A3B2	170.78±20.57ab	242.42±24.07ab	531.00±52.34a	944.21±76.57ab	18.08±1.43b	25.74±2.50ab	56.18±1.59ab
	A3B3	230.42±23.86a	266.92±35.41ab	345.68±62.44b	843.03±70.49b	27.31±1.04a	31.84±4.92ab	40.85±5.25c
	η²_A	0.395	0.542	0.289	0.572	0.084	0.067	0.019
	η²_B	0.914	0.902	0.809	0.877	0.938	0.615	0.883
	η²_A×B	0.216	0.66	0.115	0.156	0.435	0.408	0.35